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When a flattened root is a sublayer, its ruler guides were found by checking whether the sublayer held any and then walking its whole subtree again to collect them. They are now collected in one descent through the sublayer and its sublayers, looking at each of their children once. The guides, their order and the layer each one goes to are unchanged. A test-only record of the nodes looked at checks that each one is looked at once and that the guides keep their paint order.